Considering its TES propably never.
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Hmm okay, i thought that choice was the only choice in the game which stopped progression of another quest.
Come to think of it, when i did the first quest for the Stormcloaks, it came up with "Failed: Join the empire", its probable that i can still do the quests, just not join the fight.
Anyway i just think its a bit silly that i'm doing stuff for the Jarl in Solitude while being part of the Stormcloaks.
Actually I think it does block you off, I got to a point where I had to make a decision whether I want to Join the Stormcloaks and it seemed like it would put me in ill favor with the imperials, so since I hadn't done their quests yet I put a hold on joining them.
You're only susceptible to silver when in Beast Form, wearing silver armour or using silver weapons does not affect you either. The only disadvantages of being a werewolf are when you in beast mode and can't sneak and loot etc.
Some NPC's will say you smell like a dog though :D
Does anyone play in 3rd person view? It looks so much more awesome, but it's just crap to control in my opinion